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TNF- can induce the proliferation of fibroblast-like synoviocytes (FLS) and synovial cells by activating nuclear factor kappa-B (NF-B) and extracellular regulated protein kinases (Erk)-1/2-E26 transformation-specific (ETS)-1 signaling pathway, respectively (70, 71), resulting in the secretion of a variety of inflammatory mediators like IL-6, matrix metalloproteinases (MMP)-1, and MMP-3 to increase inflammation (72)
Funding This work was supported by the National Major Scientific and Technological Special Project for Significant New Drugs Development during the Thirteenth Five-year Plan Period (Grant number: 2020ZX09201-003), National Natural Science Foundation of China (Grant number: 32000985), Key Disciplines Construction Program of Traditional Chinese Medicine (Integrated Traditional Chinese and Western Medicine) during the Thirteenth Five-year Plan Period of Zhejiang province (Grant number: 2017-XK-A35), Zhejiang Provincial Natural Science Foundation (Grant number: LQ21H300003), Zhejiang Province Medical and Health Science Research Project (Grant number: 2021KY666), and Zhejiang Pharmaceutical Association (Grant number: 2019ZYY12)
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